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I presume that you want to know the difference between a notifier and an autoresponder, is that correct?
The notifier is for when you want to send an email notification to a specific email address when a form is submitted and an autoresponder is for when you want to provide your submitters an automated response for when they fill your form.
Here: http://www.jotform.com/help/25-Setting-Up-Email-Notifications you can find more information about a notifier and here: http://www.jotform.com/help/26-Creating-a-Form-Autoresponder about an autoresponder.
Let us know if you've further questions.